ftdnastat Русский

Ukraine: ancient DNA

207
Y-DNA
samples
82
mtDNA
samples

People whose remains archaeologists found and whose DNA labs have read. The date is the middle of the dating interval, radiocarbon or by archaeological context. Branches link to branch pages where one exists. All archaeological cultures

Y-DNA

By period #

35,000–34,000 BCE
1 · 0.5%
9,000–8,000 BCE
10 · 4.8%
8,000–7,000 BCE
1 · 0.5%
7,000–6,000 BCE
4 · 1.9%
6,000–5,000 BCE
45 · 21.7%
5,000–4,000 BCE
16 · 7.7%
4,000–3,000 BCE
29 · 14.0%
3,000–2,000 BCE
11 · 5.3%
2,000–1,000 BCE
11 · 5.3%
1,000–0 BCE
46 · 22.2%
0–1,000 CE
11 · 5.3%
1,000–2,000 CE
22 · 10.6%
Y-DNA

Branches #

I-PF6902
21 · 10.1%
R-PF6287
18 · 8.7%
I-L38
12 · 5.8%
R-Z2106
10 · 4.8%
I-L1286
6 · 2.9%
G-BY28175
6 · 2.9%
R-PF6162
5 · 2.4%
E-V13
5 · 2.4%
R-YP4141
4 · 1.9%
G-PF3177
4 · 1.9%
R-Z2109
4 · 1.9%
Q-M346
3 · 1.4%
Y-DNA

Cultures #

All archaeological cultures

mtDNA

By period #

9,000–8,000 BCE
2 · 2.4%
8,000–7,000 BCE
1 · 1.2%
6,000–5,000 BCE
1 · 1.2%
4,000–3,000 BCE
9 · 11.0%
3,000–2,000 BCE
11 · 13.4%
2,000–1,000 BCE
2 · 2.4%
1,000–0 BCE
28 · 34.1%
0–1,000 CE
10 · 12.2%
1,000–2,000 CE
12 · 14.6%
mtDNA

Branches #

U5b2
3 · 3.7%
J1c2
3 · 3.7%
N1a1a1a
2 · 2.4%
T2b
2 · 2.4%
U5a2b
2 · 2.4%
U4c1
2 · 2.4%
X
2 · 2.4%
J2b1a
2 · 2.4%
A
2 · 2.4%
H1
2 · 2.4%
C4a1a
2 · 2.4%
U4
1 · 1.2%

Paternal lines of the ancients #

Source: FamilyTreeDNA Discover.

Paternal lines of the ancients
Sample Culture Site Date Branch Study
Buran Kaya 3C BuranKaya3C Gravettian Buran-Kaya 3, Crimea 35,000 BCE F-M89 Bennett et al. 2023
Vasilevka 545 NEO545 Mesolithic Hunter-Gatherer Vasilevka-I, Dnepr valley 8,879 BCE I-FT344596 ? Allentoft et al. 2024
Vasylivka 1819 I1819 Mesolithic Hunter-Gatherer Vasylivka, Zaporizhia 8,708 BCE R-CTS7500 Akbari et al. 2026; Allen Ancient Genome Diversity Project; Jones et al. 2017; Mathieson et al. 2018
Vasilevka 492 NEO492 Mesolithic Hunter-Gatherer Vasilevka-I, Dnepr valley 8,705 BCE I-P37 ? Allentoft et al. 2024
Vasilevka 497 NEO497 Mesolithic Hunter-Gatherer Vasilevka-I, Dnepr valley 8,705 BCE R-M420 ? Allentoft et al. 2024
Vasilevka 501 NEO501 Mesolithic Hunter-Gatherer Vasilevka-I, Dnepr valley 8,672 BCE Q-M346 ? Allentoft et al. 2024
Vasilevka 549 NEO549 Mesolithic Hunter-Gatherer Vasilevka-I, Dnepr valley 8,615 BCE I-S21204 ? Allentoft et al. 2024
Deriivka 125 ukr125 Mesolithic Hunter-Gatherer Deriivka, Kirovohrad Oblast 8,599 BCE I-FT344596 Mattila et al. 2023
Vasilievka 2 VSL002 Mesolithic Hunter-Gatherer Vasilievka I, Dnepr 8,465 BCE I-S21825 ? Posth et al. 2023
Vasilevka 4 VSL004 Mesolithic Hunter-Gatherer Vasilevka, Dnepr 8,465 BCE R-PF6287 ? Posth et al. 2023
Vasylivka 1763 I1763 Mesolithic Hunter-Gatherer Vasylivka, Zaporizhia 8,131 BCE I-FT344596 Akbari et al. 2026; Allen Ancient Genome Diversity Project; Mathieson et al. 2018
Saint Petersburg 7 I1734 Mesolithic Hunter-Gatherer Vasylivka, Zaporizhia 7,209 BCE R-PF6362 Allen Ancient Genome Diversity Project; Mathieson et al. 2018; Rohland et al. 2022
Deriivka 5876 I5876 Mesolithic Hunter-Gatherer Deriivka I, Kirovohrad Oblast 6,882 BCE R-YP5056 Mathieson et al. 2018
Deriivka 111 ukr111 Mesolithic Hunter-Gatherer Deriivka, Kirovohrad Oblast 6,676 BCE I-L703 Mattila et al. 2023
Mamay-Gora 8 UKR008 Neolithic Steppe Mamay-Gora, Velyka Znamianka, Vasylivsky, Zaporizhzhia 6,500 BCE R-PF6343 ? Saag et al. 2025
Deriivka 102 ukr102 Mesolithic Hunter-Gatherer Deriivka, Kirovohrad Oblast 6,435 BCE R-YP5056 Mattila et al. 2023
Deriivka 108 ukr108 Mesolithic Hunter-Gatherer Deriivka, Kirovohrad Oblast 5,993 BCE Q-FTF30 Mattila et al. 2023
Vil’nyanka 3715 I3715 Mariupol Vil’nyanka, Zaporizhia Oblast 5,576 BCE I-L703 Akbari et al. 2026; Mathieson et al. 2018
Igren 1 IGR001 Serednii Stih Igren’ 8, Dnipro 5,565 BCE Q-M346 ? Posth et al. 2023
Yasynuvatka 27994 I27994 Bug–Dniester Yasynuvatka, Vilnianskyi, Zaporizhzhia 5,557 BCE I-FTA13672 ? Nikitin et al. 2025
Vovnigi 2-1 VO2001 Neolithic Eastern European Vovnigi II, Solonyansky district 5,490 BCE R-V88 ? Posth et al. 2023
Vil’nyanka 5869 I5869 Mariupol Vil’nyanka, Zaporizhia Oblast 5,489 BCE I-P215 ? Mathieson et al. 2018
Vil’nyanka 5870 I5870 Mariupol Vil’nyanka, Zaporizhia Oblast 5,460 BCE I-L703 Akbari et al. 2026; Mathieson et al. 2018
Vovnigi 5 VO1005 Neolithic Eastern European Vovnigi I, Solonyansky district 5,443 BCE I-P222 ? Posth et al. 2023
Vovnigi 3 VO1003 Neolithic Eastern European Vovnigi I, Solonyansky district 5,443 BCE I-S11321 ? Posth et al. 2023
Vovnigi 1 VO1001 Neolithic Eastern European Vovnigi I, Solonyansky district 5,443 BCE I-S2599 ? Posth et al. 2023
Vovnigi 4 VO1004 Neolithic Eastern European Vovnigi I, Solonyansky district 5,443 BCE I-Y8551 ? Posth et al. 2023
Vil’nyanka 5872 I5872 Mariupol Vil’nyanka, Zaporizhia Oblast 5,424 BCE I-L703 Akbari et al. 2026; Mathieson et al. 2018
Deriivka 4112 I4112 Mariupol Deriivka I, Kirovohrad Oblast 5,403 BCE R-Y61088 Lazaridis et al. 2025; Mathieson et al. 2018
Deriivka 4114 I4114 Mariupol Deriivka I, Kirovohrad Oblast 5,401 BCE R-Y61088 Mathieson et al. 2018
Deriivka 5888 I5888 Mariupol Deriivka I, Kirovohrad Oblast 5,397 BCE I-L703 Lazaridis et al. 2025; Mathieson et al. 2018
Vil’nyanka 3716 I3716 Mariupol Vil’nyanka, Zaporizhia Oblast 5,397 BCE I-Y8551 Akbari et al. 2026; Mathieson et al. 2018
Saint Petersburg 4 I1738 Dnieper–Donets Vovnigi, Dnipropetrovsk Oblast 5,360 BCE I-Y8551 Akbari et al. 2026; Allen Ancient Genome Diversity Project; Mathieson et al. 2018
Mariupol 31730 I31730 Bug–Dniester Mariupol, Mariupol, Donetsk 5,355 BCE I-Y8551 ? Nikitin et al. 2025
Deriivka 5 ukr005 Neolithic Eastern European Deriivka, Kirovohrad Oblast 5,346 BCE R-Y61088 Mattila et al. 2023
Mamaj Gora 270 NEO270 Cucuteni–Trypillia Mamaj Gora, North Pontic Steppe 5,343 BCE I-FTA13672 ? Allentoft et al. 2024
Vil’nyanka 5957 I5957 Mariupol Vil’nyanka, Zaporizhia Oblast 5,337 BCE I-P222 ? Mathieson et al. 2018
Mamaj Gora 268 NEO268 Cucuteni–Trypillia Mamaj Gora, North Pontic Steppe 5,295 BCE R-PF6287 ? Allentoft et al. 2024
Deriivka 5893 I5893 Mariupol Deriivka I, Kirovohrad Oblast 5,294 BCE R-Y61088 Lazaridis et al. 2025; Mathieson et al. 2018
Deriivka 3717 I3717 Mariupol Deriivka I, Kirovohrad Oblast 5,292 BCE I-L702 ? Mathieson et al. 2018
Yasinovatka 160 ukr160 Neolithic Eastern European Yasinovatka, Zaporizhia Oblast 5,285 BCE I-FTA13672 ? Mattila et al. 2023
Deriivka 3718 I3718 Mariupol Deriivka I, Kirovohrad Oblast 5,280 BCE R-Y61088 Mathieson et al. 2018
Deriivka 5879 I5879 Mariupol Deriivka I, Kirovohrad Oblast 5,265 BCE R-Y61088 ? Mathieson et al. 2018
Deriivka 5886 I5886 Mariupol Deriivka I, Kirovohrad Oblast 5,263 BCE I-L703 Lazaridis et al. 2025; Mathieson et al. 2018
Deriivka 5890 I5890 Mariupol Deriivka I, Kirovohrad Oblast 5,233 BCE R-Y61088 Mathieson et al. 2018; Mattila et al. 2023
Yasinovatka 158 ukr158 Neolithic Eastern European Yasinovatka, Zaporizhia Oblast 5,195 BCE R-PF6287 ? Mattila et al. 2023
Deriivka 5878 I5878 Mariupol Deriivka I, Kirovohrad Oblast 5,184 BCE R-Y61088 ? Lazaridis et al. 2025; Mathieson et al. 2018
Deriivka 117 ukr117 Neolithic Eastern European Deriivka, Kirovohrad Oblast 5,182 BCE I-L703 ? Mattila et al. 2023
Yasynuvatka 27990 I27990 Bug–Dniester Yasynuvatka, Vilnianskyi, Zaporizhzhia 5,155 BCE Q-M242 ? Nikitin et al. 2025
Vil’nyanka 3714 I3714 Mariupol Vil’nyanka, Zaporizhia Oblast 5,150 BCE I-M223 ? Mathieson et al. 2018
Igren 516 NEO516 Cucuteni–Trypillia Igren, Dnepr valley 5,146 BCE Q-M346 ? Allentoft et al. 2024
Deriivka 5875 I5875 Mariupol Deriivka I, Kirovohrad Oblast 5,145 BCE I-L703 ? Mathieson et al. 2018
Deriivka 5881 I5881 Mariupol Deriivka I, Kirovohrad Oblast 5,133 BCE R-Y61088 Lazaridis et al. 2025; Mathieson et al. 2018
Deriivka 5883 I5883 Mariupol Deriivka I, Kirovohrad Oblast 5,128 BCE R-Y64721 ? Mathieson et al. 2018
Yasinovatka 161 ukr161 Neolithic Eastern European Yasinovatka, Zaporizhia Oblast 5,107 BCE I-FTA13672 ? Mattila et al. 2023
Vovnigi 498 NEO498 Neolithic Eastern European Vovnigi II, Dnepr valley 5,106 BCE R-PF6287 Allentoft et al. 2024
Yasinovatka 159 ukr159 Neolithic Eastern European Yasinovatka, Zaporizhia Oblast 5,105 BCE I-FTA13672 ? Mattila et al. 2023
Nikol’ske 144 ukr144 Neolithic Eastern European Nikol’ske, Dnipropetrovsk Oblast 5,078 BCE I-S22311 ? Mattila et al. 2023
Volnensky 304 NEO304 Cucuteni–Trypillia Volnensky, Dnepr valley 5,056 BCE I-L703 Allentoft et al. 2024
Vovnigi 524 NEO524 Cucuteni–Trypillia Vovnigi II, Dnepr valley 5,026 BCE I-L702 ? Allentoft et al. 2024
Vovnigi 552 NEO552 Bug–Dniester Vovnigi I, Dnepr valley 5,020 BCE I-FTA13672 ? Allentoft et al. 2024
Vovnigi 507 NEO507 Cucuteni–Trypillia Vovnigi II, Dnepr valley 4,937 BCE R-PF6287 ? Allentoft et al. 2024
Vovnigi 528 NEO528 Cucuteni–Trypillia Vovnigi I, Dnepr valley 4,892 BCE I-FTA13672 ? Allentoft et al. 2024
Deriivka 3719 I3719 Mariupol Deriivka I, Kirovohrad Oblast 4,889 BCE I-FTH81 Lazaridis et al. 2025; Mathieson et al. 2018
Volnensky 302 NEO302 Cucuteni–Trypillia Volnensky, Dnepr valley 4,889 BCE I-Y8551 Allentoft et al. 2024
Vovnigi 514 NEO514 Cucuteni–Trypillia Vovnigi II, Dnepr valley 4,673 BCE I-FTA13672 ? Allentoft et al. 2024
Nikol’ske 147 ukr147 Neolithic Eastern European Nikol’ske, Dnipropetrovsk Oblast 4,668 BCE I-L703 ? Mattila et al. 2023
Yasinovatka 162 ukr162 Neolithic Eastern European Yasinovatka, Zaporizhia Oblast 4,668 BCE Q-FTF30 Mattila et al. 2023
Vovnigi 502 NEO502 Cucuteni–Trypillia Vovnigi II, Dnepr valley 4,666 BCE I-FTE66752 Allentoft et al. 2024
Vasilevsky 305 NEO305 Cucuteni–Trypillia Vasilevsky, Dnepr valley 4,661 BCE I-Y8551 Allentoft et al. 2024
Nikol’ske 33 ukr033 Neolithic Eastern European Nikol’ske, Dnipropetrovsk Oblast 4,572 BCE I-S2599 ? Mattila et al. 2023
Lysa Gora 262 NEO262 Cucuteni–Trypillia Lysa Gora, North Pontic Steppe 4,563 BCE R-Y61088 Allentoft et al. 2024
Deriivka 27283 I27283 Serednii Stih Deriivka II, Onufryiv, Kirovohrad 4,353 BCE I-L703 ? Nikitin et al. 2025
Igren 27930 I27930 Serednii Stih Igren-8, Dnipro, Dnipropetrovsk 4,200 BCE R-L761 ? Nikitin et al. 2025
Deriivka 27282 I27282 Serednii Stih Deriivka II, Onufryiv, Kirovohrad 4,182 BCE I-L703 ? Nikitin et al. 2025
Vynohradne 1430 I1430 Serednii Stih Vynohradne, Zaporizhzhia 4,096 BCE I-Y8551 ? Nikitin et al. 2025
Kartal 6 KTL006 Cernavodă Kartal, Odessa Oblast 4,058 BCE I-PF6902 ? Penske et al. 2023
Verteba 3151 I3151 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,800 BCE E-M35 ? Mathieson et al. 2018; Nikitin et al. 2025
Majaky 3 MAJ003 Usatove Majaky, Odessa Oblast 3,777 BCE G-FTE180 Penske et al. 2023
Verteba Cave 2110 I2110 Cucuteni–Trypillia Verteba Cave, Chortkiv, Ternopil 3,747 BCE G-P303 ? Mathieson et al. 2018; Nikitin et al. 2025
Verteba 106 VERT106 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,720 BCE G-FTE180 Gelabert et al. 2022 (Sci. Rep.)
Verteba 33 VERT033 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,720 BCE I-FTK242 Gelabert et al. 2022 (Sci. Rep.)
Verteba Cave 7923 I7923 Cucuteni–Trypillia Verteba Cave, Chortkiv, Ternopil 3,719 BCE J-CTS7683 ? Nikitin et al. 2025
Molyukhiv-Bugor 1424 I1424 Serednii Stih Molyukhiv-Bugor, Chygyryn, Cherkasy 3,712 BCE I-PF6902 ? Nikitin et al. 2025
Verteba 117 VERT117 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,710 BCE I-BY160929 Gelabert et al. 2022 (Sci. Rep.)
Verteba 1926 I1926 Cucuteni–Trypillia Verteba Cave, Chortkiv, Ternopil 3,708 BCE G-FTE180 Mathieson et al. 2018; Nikitin et al. 2025
Kartak 5 KTL005 Cernavodă Kartal, Odessa Oblast 3,707 BCE R-BY15339 ? Penske et al. 2023
Verteba 103 VERT103 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE C-FTJ33661 Gelabert et al. 2022 (Sci. Rep.)
Verteba 31 VERT031 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE G-BY201413 Gelabert et al. 2022 (Sci. Rep.)
Verteba 100 VERT100 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE G-FTA29810 Gelabert et al. 2022 (Sci. Rep.)
Verteba 30 VERT030 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE G-FTE180 Gelabert et al. 2022 (Sci. Rep.)
Verteba 105 VERT105 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE G-FTE180 Gelabert et al. 2022 (Sci. Rep.)
Verteba 104 VERT104 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE G-FTF153 Gelabert et al. 2022 (Sci. Rep.)
Verteba 111 VERT111 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE G-FTH126 ? Gelabert et al. 2022 (Sci. Rep.)
Verteba 29 VERT029 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE G-Y240410 Gelabert et al. 2022 (Sci. Rep.)
Verteba 107 VERT107 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE G-Z39358 Gelabert et al. 2022 (Sci. Rep.)
Verteba 28 VERT028 Cucuteni–Trypillia Verteba Cave, Ternopil Oblast 3,705 BCE I-BY136773 Gelabert et al. 2022 (Sci. Rep.)
Revova 7929 I7929 Usatove Revova, Berezivka, Odesa 3,675 BCE J-FT265251 ? Nikitin et al. 2025
Verteba Cave 2111 I2111 Cucuteni–Trypillia Verteba Cave, Bilche-Zolote, Chortkiv, Ternopil 3,672 BCE G-P15 ? Mathieson et al. 2018; Nikitin et al. 2025
Mayaky 1423 I1423 Usatove Mayaky, Belyaiivsky, Odesa 3,654 BCE R-M459 Nikitin et al. 2025
Usatove 6 UBK006 Usatove Usatove-Bolšoj Kujalnik, Odessa Oblast 3,618 BCE I-FTK242 Penske et al. 2023
Verteba Cave 13064 I13064 Cucuteni–Trypillia Verteba Cave, Chortkiv, Ternopil 3,588 BCE I-FTH75 Nikitin et al. 2025
Lysa Gora 265 NEO265 Cucuteni–Trypillia Lysa Gora, North Pontic Steppe 3,585 BCE I-L702 ? Allentoft et al. 2024
Оleksandria 6558 I6558 Serednii Stih Оleksandria, Kharkiv Region, Kupianskyi District 3,504 BCE I-P222 ? Lazaridis et al. 2025
Mayaky 12704 I12704 Usatove Mayaky, Belyaiivsky, Odesa 3,500 BCE E-M78 ? Nikitin et al. 2025
Kartal 1 KTL001 Cernavodă Kartal, Odessa Oblast 3,423 BCE I-S12195 ? Penske et al. 2023
Katarzhyno 12168 I12168 Yamnaya Katarzhyno kurgan group, Znamianka, Ivanovsky 2,964 BCE R-Z2109 ? Nikitin et al. 2025
Ilyatka 1 ILK001 Globular Amphora Ilyatka, Staraya Sinyava, Khmelnytskyi Oblast 2,823 BCE I-Y167771 Mathieson et al. 2018
Majaky 17 MAJ017 Usatove Majaky, Odessa Oblast 2,808 BCE R-M12149 ? Penske et al. 2023
Ilyatka 2 ILK002 Globular Amphora Ilyatka, Staraya Sinyava, Khmelnytskyi Oblast 2,784 BCE I-Y167771 Mathieson et al. 2018
Deriivka 5884 I5884 Mariupol Deriivka I, Kirovohrad Oblast 2,783 BCE R-BY44400 ? Lazaridis et al. 2025; Mathieson et al. 2018
Kam’yana Mohyla 20975 I20975 Yamnaya Kam’yana Mohyla, Myrne, Zaporizhzhia 2,749 BCE R-BY132259 ? Nikitin et al. 2025
Sychavka 12843 I12843 Yamnaya Sychavka, Odesa 2,679 BCE R-BY44400 ? Nikitin et al. 2025
Vapnyarka 12619 I12619 Yamnaya Vapnyarka, Odesa 2,665 BCE R-Z2109 ? Nikitin et al. 2025
Katarzhyno 11999 I11999 Yamnaya Katarzhyno kurgan group, Znamianka, Ivanovsky 2,551 BCE R-Z2109 ? Nikitin et al. 2025
Liubasha 16674 I16674 Multi-cordoned Ware Liubasha, Novohryhorivka, Mykolaivskyi 2,188 BCE R-BY44400 ? Nikitin et al. 2025
Igren 509 NEO509 Neolithic Eastern European Igren, Dnepr valley 2,114 BCE R-M12149 ? Allentoft et al. 2024
Liubasha 7925 I7925 Multi-cordoned Ware Liubasha, Novohryhorivka, Mykolaivskyi 1,871 BCE R-Y86480 ? Nikitin et al. 2025
Sychavka 12231 I12231 Multi-cordoned Ware Sychavka, Lymansky, Odesa 1,841 BCE R-KMS75 ? Nikitin et al. 2025
Beremiany 643 poz643 Komarów Beremiany, Tarnopil Province 1,810 BCE I-CTS10937 ? Chyleński et al. 2023
Aleksandria 6561 I6561 Srubnaya Aleksandria, Kupyansk, Kharkiv Oblast 1,700 BCE R-Z93 ? Mathieson et al. 2018
Biały Potok 296 poz296 Komarów Biały Potok, Tarnopil province 1,531 BCE R-Z92 ? Chyleński et al. 2023
Liubasha 12235 I12235 Multi-cordoned Ware Liubasha, Novohryhorivka, Mykolaivskyi 1,498 BCE R-PF6162 ? Nikitin et al. 2025
Mykolaiv 31 MJ-31 Cimmerian Dykyi Sad hillfort, "Citadel", Mykolaiv city, Mykolaiv region 1,186 BCE R-M417 ? Järve et al. 2019
Petrykiv 171 UKR171 Early Vysotska Petrykiv, Petrykiv, Ternopil, Ternopil 1,166 BCE R-PF6162 ? Saag et al. 2025
Dykyi Sad 150 UKR150 Bilozerska Dykyi Sad, Mykolaiv 1,100 BCE R-PF6162 ? Saag et al. 2025
Kumy 66 UKR066 Cimmerian Kumy, Krasnograd, Kharkiv 1,057 BCE Q-YP3953 ? Saag et al. 2025
Petrykiv 170 UKR170 Early Vysotska Petrykiv, Petrykiv, Ternopil, Ternopil 1,050 BCE R-M459 ? Saag et al. 2025
Kartal 7 UKR007 Hallstatt Kartal, Orlivka, Izmail, Odesa 913 BCE E-CTS8814 ? Saag et al. 2025
Kartal 2 UKR002 Hallstatt Kartal, Orlivka, Izmail, Odesa 800 BCE C-Z44530 ? Saag et al. 2025
Kartal 5 UKR005 Hallstatt Kartal, Orlivka, Izmail, Odesa 800 BCE G-PF3239 ? Saag et al. 2025
Kupievakha 105 UKR105 Scythian Kupievakha, Kupevaha, Bohodukhiv, Kharkiv 675 BCE R-FTA11514 Saag et al. 2025
Cheremushna 111 UKR111 Scythian Cheremushna, Cheremushna, Bohoduhiv, Kharkiv 657 BCE R-YP5049 Saag et al. 2025
Medvyn 35 UKR035 Scythian Medvyn, Bila Tserkva, Kyiv 650 BCE R-FTD71214 Saag et al. 2025
Medvyn 39 UKR039 Scythian Medvyn, Bila Tserkva, Kyiv 650 BCE R-M420 ? Saag et al. 2025
Blagoveschenka 30 AS30 Scythian Blagoveschenka, Zaporizhia 647 BCE Q-YP3953 ? Andreeva et al. 2025
Karavan 116 UKR116 Scythian Karavan, Karavan, Lyubotyn, Kharkiv 645 BCE R-Z282 ? Saag et al. 2025
Medwin 14 I27512 Scythian Gorchakov's Wood, Medwin village, Boguslav district, Kiev region 643 BCE R-FTD71214 Akbari et al. 2026; Järve et al. 2019
Scythian 9 scy009 Western Scythian Kurgan 1, Starosillya, Odessa Oblast 618 BCE R-FTA9989 Krzewińska et al. 2018 (Sci. Adv.)
Verteba 1927 I1927 Scythian Verteba Cave, Bilche-Zolote, Ternopil Oblast 608 BCE G-Y11074 ? Akbari et al. 2026
Medwin 33 MJ-33 Scythian Gorchakov's Wood, Medwin village, Boguslav district, Kiev region 603 BCE R-PF6162 ? Järve et al. 2019
Mala Rogozianka 113 UKR113 Scythian Mala Rogozianka, Mala Rogozianka, Bohoduhiv, Kharkiv 600 BCE Q-L332 ? Saag et al. 2025
Medvyn 36 UKR036 Scythian Medvyn, Bila Tserkva, Kyiv 599 BCE R-M458 ? Saag et al. 2025
Medwin 16 I27505 Scythian Gorchakov's Wood, Medwin village, Boguslav district, Kiev region 594 BCE J-Z7706 ? Akbari et al. 2026; Järve et al. 2019
Oleksandrivskyi 153 UKR153 Scythian Oleksandrivskyi necropolis, Naberezhna Street, Mykolaiv City 573 BCE R-CTS1211 ? Saag et al. 2025
Kup'evaha 35 MJ-35 Scythian Kup'evaha, Bogodukhov district, Kharkiv region 565 BCE Q-YP771 ? Järve et al. 2019
Bilsk Hillfort 91 UKR091 Scythian Bilsk hillfort, Bilsk, Poltava, Poltava 450 BCE E-BY179384 Saag et al. 2025
Grishkivka 104 UKR104 Scythian Grishkivka, Grishkivka, Zmiiv, Kharkiv 425 BCE R-FTH5039 ? Saag et al. 2025
Bilsk Hillfort 89 UKR089 Scythian Bilsk hillfort, Bilsk, Poltava, Poltava 400 BCE E-BY179384 Saag et al. 2025
Chersonesus 21475 CGG021475 Iron Age Crimean Chersonesus City, Crimea 400 BCE E-BY5022 ? McColl et al. 2024 (bioRxiv)
Chersonesus 21473 CGG021473 Iron Age Crimean Chersonesus City, Crimea 400 BCE E-BY6062 McColl et al. 2024 (bioRxiv)
Pisochyn 133 UKR133 Scythian Pisochyn, Pisochyn, Kharkiv, Kharkiv 400 BCE R-FTD77954 ? Saag et al. 2025
Pisochyn 131 UKR131 Scythian Pisochyn, Pisochyn, Kharkiv, Kharkiv 400 BCE R-FTH50000 Saag et al. 2025
Pisochyn 132 UKR132 Scythian Pisochyn, Pisochyn, Kharkiv, Kharkiv 400 BCE R-FTH50000 Saag et al. 2025
Bilsk Hillfort 83 UKR083 Scythian Bilsk hillfort, Bilsk, Poltava, Poltava 400 BCE R-M198 ? Saag et al. 2025
Akkermen 28 AS28 Scythian Akkermen, Odesa 400 BCE R-M417 ? Andreeva et al. 2025
Bilsk Hillfort 90 UKR090 Scythian Bilsk hillfort, Bilsk, Poltava, Poltava 350 BCE E-BY179384 ? Saag et al. 2025
Gajmanovo Pole 34 AS34 Scythian Gajmanovo Pole, Zaporizhia 350 BCE G-FT34 ? Andreeva et al. 2025
Gajmanovo Pole 33 AS33 Scythian Gajmanovo Pole, Zaporizhia 350 BCE J-BY70 ? Andreeva et al. 2025
Vesele 109 UKR109 Scythian Vesele, Vesele, Kharkiv, Kharkiv 350 BCE Q-BZ433 Saag et al. 2025
Kut 40 AS40 Scythian Kut, Dnipropetrovsk 350 BCE R-BY86775 ? Andreeva et al. 2025
Vesele 110 UKR110 Scythian Vesele, Vesele, Kharkiv, Kharkiv 350 BCE R-FT155201 Saag et al. 2025
Mamay-Gora 14 UKR014 Scythian Mamay-Gora, Velyka Znamianka, Vasylivsky, Zaporizhzhia 350 BCE R-S23592 ? Saag et al. 2025
Oleksandrivskyi 152 UKR152 Hellenistic Oleksandrivskyi necropolis, Naberezhna Street, Mykolaiv City 299 BCE E-FGC57496 ? Saag et al. 2025
Merchik 32 MJ-32 Scythian St Merchik, Krasnokutsk district, Kharkiv region 296 BCE R-M417 ? Järve et al. 2019
Merchik 15 MJ-15 Scythian Stary Merchik village, Valki district, Kharkiv region 296 BCE R-Z2124 ? Järve et al. 2019
Kolomak 95 UKR095 Scythian Kolomak, Kolomak City, Kharkiv 296 BCE R-Z2125 ? Saag et al. 2025
Kolomak 34 MJ-34 Scythian Kolomak hillfort, Kolomak city, Kharkiv region 281 BCE R-Z94 ? Järve et al. 2019
Chersonesus 21478 CGG021478 Iron Age Crimean Chersonesus City, Crimea 245 BCE L-FTG91317 McColl et al. 2024 (bioRxiv)
Belyaus 7 AS7 Late Scythian Belyaus, Crimea 229 BCE J-BY22491 ? Andreeva et al. 2025
Igren 518 NEO518 Iron Age Eastern European Igren, Dnepr valley 202 BCE R-S23592 ? Allentoft et al. 2024
Yevpatoria 13 AS13 Late Scythian Yevpatoria, Crimea 125 BCE I-CTS10228 ? Andreeva et al. 2025
Belyaus 11 AS11 Late Scythian Belyaus, Crimea 125 BCE J-M410 ? Andreeva et al. 2025
Kermenchik 1 AS1 Late Scythian Neapol Scythian, Crimea 30 BCE N-BY76278 ? Andreeva et al. 2025
Shyshaky 126 UKR126 Chernyakhіv Shуshakу, Shyshaky, Shyshaky, Poltava 228 CE R-YP6082 Saag et al. 2025
Kerch 1 KER_1 Hun Kerch, Crimea 346 CE J-BY116167 ? Veeramah et al. 2018
Zolochiv 102 UKR102 Chernyakhіv Zolochiv, Chalogo street, Zolochiv City, Bohodukhiv, Kharkiv 350 CE E-V13 ? Saag et al. 2025
Shyshaky 123 UKR123 Chernyakhіv Shуshakу, Shyshaky, Shyshaky, Poltava 350 CE R-Z2125 ? Saag et al. 2025
Suvlu-Khaye 17683 CGG017683 Gothic Suvlu-Khaye, Crimea 550 CE G-FTB38820 McColl et al. 2024 (bioRxiv)
Suvlu-Khaye 17682 CGG017682 Sarmatian Suvlu-Khaye, Crimea 550 CE R-FTH7422 McColl et al. 2024 (bioRxiv)
Bochkove 147 UKR147 Saltiv Bochkove, Bochkove, Chuhuev, Kharkiv 783 CE R-BY60213 ? Saag et al. 2025
Verkhnii Saltiv 137 UKR137 Saltiv Verkhnii Saltiv, Verkhnii Saltiv, Chuhuev, Kharkiv 850 CE C-Y11606 ? Saag et al. 2025
Verkhnii Saltiv 135 UKR135 Saltiv Verkhnii Saltiv, Verkhnii Saltiv, Chuhuev, Kharkiv 850 CE R-FTD77954 ? Saag et al. 2025
Verkhnii Saltiv 136 UKR136 Saltiv Verkhnii Saltiv, Verkhnii Saltiv, Chuhuev, Kharkiv 850 CE R-FTD77954 ? Saag et al. 2025
Verkhnii Saltiv 138 UKR138 Saltiv Verkhnii Saltiv, Verkhnii Saltiv, Chuhuev, Kharkiv 850 CE R-FTD77954 ? Saag et al. 2025
Shestovitsa 539 VK539 Viking Age Ukrainian Shestovitsa 1,050 CE I-BY65928 Margaryan et al. 2020
Chernigov 542 VK542 Viking Age Ukrainian Chernigov 1,050 CE I-S20602 ? Margaryan et al. 2020
Pidhirtsi 5 PDH005 East Slavic Pidhirtsi (Podhorce), Lviv Oblast 1,150 CE E-V13 ? Gretzinger et al. 2025
Pidhirtsi 4 PDH004 East Slavic Pidhirtsi (Podhorce), Lviv Oblast 1,150 CE I-CTS10228 ? Gretzinger et al. 2025
Pidhirtsi 7 PDH007 East Slavic Pidhirtsi (Podhorce), Lviv Oblast 1,150 CE I-CTS10228 ? Gretzinger et al. 2025
Pidhirtsi 9 PDH009 East Slavic Pidhirtsi (Podhorce), Lviv Oblast 1,150 CE I-PF3892 ? Gretzinger et al. 2025
Pidhirtsi 8 PDH008 East Slavic Pidhirtsi (Podhorce), Lviv Oblast 1,150 CE I-S20602 ? Gretzinger et al. 2025
Pidhirtsi 3 PDH003 East Slavic Pidhirtsi (Podhorce), Lviv Oblast 1,150 CE J-Z30685 ? Gretzinger et al. 2025
Velyka Znamianka 27 UKR027 Cuman Velyka Znamianka, Velyka Znamianka, Vasylivsky, Zaporizhzhia 1,150 CE N-PH1896 ? Saag et al. 2025
Pidhirtsi 6 PDH006 East Slavic Pidhirtsi (Podhorce), Lviv Oblast 1,150 CE R-L1029 ? Gretzinger et al. 2025
Pidhirtsi 12 PDH012 East Slavic Pidhirtsi (Podhorce), Lviv Oblast 1,150 CE R-Y33 ? Gretzinger et al. 2025
Korolówka 2 KRW002 Old Ruthenian Korolówka, Ternopil Oblast 1,250 CE E-A7136 ? Gretzinger et al. 2025
Korolówka 5 KRW005 Old Ruthenian Korolówka, Ternopil Oblast 1,250 CE G-FT34 ? Gretzinger et al. 2025
Korolówka 8 KRW008 Old Ruthenian Korolówka, Ternopil Oblast 1,250 CE R-FT14542 ? Gretzinger et al. 2025
Lutsk 541 VK541 Medieval Eastern European Lutsk, Volyn Oblast 1,250 CE R-FTJ67636 Margaryan et al. 2020
Mamay-Gora 12 UKR012 Cuman Mamay-Gora, Velyka Znamianka, Vasylivsky, Zaporizhzhia 1,265 CE R-PF6162 ? Saag et al. 2025
spb004 Ruthenian Kyiv 1,300 CE R-BY104638 PRJEB63275
Donets Hillfort 69 UKR069 Golden Horde Donets hillfort, Kharkiv, Novobavarsky, Kharkiv 1,300 CE R-FTD7792 Saag et al. 2025
Mamay-Gora 17 UKR017 Nogai Mamay-Gora, Velyka Znamianka, Vasylivsky, Zaporizhzhia 1,450 CE C-MPB401 Saag et al. 2025
Mamay-Gora 24 UKR024 Nogai Mamay-Gora, Velyka Znamianka, Vasylivsky, Zaporizhzhia 1,450 CE G-Y47291 ? Saag et al. 2025
Mamay-Gora 20 UKR020 Nogai Mamay-Gora, Velyka Znamianka, Vasylivsky, Zaporizhzhia 1,450 CE J-FGC4309 ? Saag et al. 2025
Mamay-Gora 16 UKR016 Nogai Mamay-Gora, Velyka Znamianka, Vasylivsky, Zaporizhzhia 1,450 CE R-L1433 ? Saag et al. 2025

Maternal lines of the ancients #

Source: YFull; no culture given.

Maternal lines of the ancients
Sample Site Date Branch
87480 Zaporiz'ka Oblast' 8,832 BCE U5b2e
87471 Zaporiz'ka Oblast' 8,121 BCE U5b2d
86636 Zaporiz'ka Oblast' 7,254 BCE U5b2d
87150 Dnipropetrovs'ka Oblast' 5,253 BCE U4
120393 Ternopil's'ka Oblast' 3,707 BCE J1c2aa
120400 Ternopil's'ka Oblast' 3,705 BCE H15a1
120420 Ternopil's'ka Oblast' 3,686 BCE N1a1a1a-a1
120390 Ternopil's'ka Oblast' 3,683 BCE U5a2-b
42461 Ternopil's'ka Oblast' 3,600 BCE HV-b
47397 Ternopil's'ka Oblast' 3,600 BCE T2b55
23691 Ternopil's'ka Oblast' 3,600 BCE H5a
48438 Vinnyts'ka Oblast' 3,275 BCE U2e1a1
40545 Ternopil's'ka Oblast' 3,250 BCE H1b1d
22948 Vinnyts'ka Oblast' 2,967 BCE W3a1a
62383 Vinnyts'ka Oblast' 2,967 BCE W3a1k
86621 Donets'ka Oblast' 2,900 BCE T1a1
102213 Donets'ka Oblast' 2,900 BCE H15b1
54340 Vinnyts'ka Oblast' 2,830 BCE U5a2b
62381 Vinnyts'ka Oblast' 2,747 BCE H2a1
56144 Vinnyts'ka Oblast' 2,740 BCE U5a1i1
56143 Vinnyts'ka Oblast' 2,711 BCE U4c1-a
62380 Vinnyts'ka Oblast' 2,667 BCE X4a1
62384 Vinnyts'ka Oblast' 2,448 BCE X4a1
62382 Vinnyts'ka Oblast' 2,034 BCE J1c2m
48437 Vinnyts'ka Oblast' 1,900 BCE J2b1a
48441 Vinnyts'ka Oblast' 1,826 BCE H1e11
145156 Odes'ka Oblast' 849 BCE U4a2a
145350 Odes'ka Oblast' 800 BCE K1a1c1a1b
144935 Odes'ka Oblast' 800 BCE U5b1a
145349 Odes'ka Oblast' 800 BCE U5a1
25597 650 BCE D4j2
32541 650 BCE J2b1a6
40169 650 BCE N1b1a-a3
144663 Kharkivs'ka Oblast' 646 BCE U2e1f1a1
151613 Kyïvs'ka Oblast' 600 BCE C4b
145355 Kiev 600 BCE U5a2b-b1
145362 Poltavs'ka Oblast' 590 BCE U2b3b1
145361 Poltavs'ka Oblast' 582 BCE H6a1b2j1c
148276 Sevastopol 470 BCE U5b1b1-b
148749 Sevastopol 470 BCE W1c
148277 Sevastopol 470 BCE U4b1b1
148750 Sevastopol 470 BCE U7b
144681 Kharkivs'ka Oblast' 400 BCE T1b
25719 350 BCE U5a1b11
44942 350 BCE J1c2
25104 350 BCE A-a1b3
145159 Zaporiz'ka Oblast' 350 BCE HV2a-a2
145364 Kharkivs'ka Oblast' 350 BCE I1a1-a
29439 300 BCE U5b2a1a2a
145354 Kiev 290 BCE T2a2
145363 Kharkivs'ka Oblast' 290 BCE T2a1a
148748 Sevastopol 276 BCE U8b1a2b5
148751 Sevastopol 264 BCE N1a1a1a
22575 200 BCE H101a
145384 Poltavs'ka Oblast' 295 CE K1a-a2
145385 Poltavs'ka Oblast' 323 CE K1c3
148725 328 CE I1i
145158 Cherkas'ka Oblast' 350 CE U4c1-a
145356 Cherkas'ka Oblast' 350 CE H2a8
145369 Poltavs'ka Oblast' 350 CE H1di
144904 Poltavs'ka Oblast' 350 CE K1b2b-a
145365 Poltavs'ka Oblast' 350 CE K1a4a1b5
148141 550 CE U4a2w1
148165 550 CE V1a10
82383 Chernihivs'ka Oblast' 1,050 CE V40a1
82188 Chernihivs'ka Oblast' 1,050 CE H5a2a
145161 Kharkivs'ka Oblast' 1,070 CE R30e1a
145160 Donets'ka Oblast' 1,250 CE A1a3
82189 Volyns'ka Oblast' 1,250 CE H1-d4a
145165 Zaporiz'ka Oblast' 1,350 CE J1c2e1
145157 Kharkivs'ka Oblast' 1,350 CE U2e1b-a2
144658 Zaporiz'ka Oblast' 1,450 CE C4a1a4a5
145351 Zaporiz'ka Oblast' 1,450 CE B4b'd'e'j
145399 Zaporiz'ka Oblast' 1,450 CE M7b1a1a1m
145353 Zaporiz'ka Oblast' 1,450 CE C4a1a4a2
144923 Zaporiz'ka Oblast' 1,450 CE U2e2a1a2
157535 Chernivets'ka Oblast' — K1a2d
120419 Ternopil's'ka Oblast' — K1a1b1-b
120394 Ternopil's'ka Oblast' — T2b81
120447 Ternopil's'ka Oblast' — H40d
120449 Ternopil's'ka Oblast' — J1c5
120418 Ternopil's'ka Oblast' — T2c1d1